Partonomy list P4, primary language: EN, subsidiary: LA, interface: EN, work in progress

mesocortex (pair)

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Top level nervous system
Level 2 encephalon Short Extended
Level 3 telencephalon Short Extended
Current level mesocortex (pair)
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Partonomy list
Short English equivalent
Short official Latin term
84039 39311 6129
mesocortex (pair)
mesocortex (par)
39314 8920
proisocortex (pair)
proisocortex (par)
39317 8921
periallocortex (pair)
periallocortex (par)
39320 8922
peripaleocortex (pair)
peripaleocortex (par)
39323 8924
periarchicortex (pair)
periarchicortex (par)
28326 12168
layers of presubiculum (pair)
strata presubiculi (par)
28431 9411
molecular layer of presubiculum (pair)
stratum moleculare presubiculi (par)
28434 9412
external principal layer of presubiculum (pair)
stratum principale externum presubiculi (par)
28437 9413
internal principal layer of presubiculum (pair)
stratum principale internum presubiculi (par)
28428 12169
layers of parasubiculum (pair)
strata parasubiculi (par)
28444 9415
molecular layer of parasubiculum (pair)
stratum moleculare parasubiculi (par)
28447 9416
cellular layer of parasubiculum (pair)
stratum cellulare parasubiculi (par)
28451 9417
layers of entorhinal cortex (pair)
strata corticis entorhinalis (par)
28454 9418
molecular layer of entorhinal cortex (pair) ; layer 1 of entorhinal cortex (pair)
stratum moleculare corticis entorhinalis (par) ; lamina 1 corticis entorhinalis (par)
28457 9421
external principal layer of entorhinal cortex (pair)
stratum principale externum corticis entorhinalis (par)
28460 9422
cell island layer of entorhinal cortex (pair); layer 2 of entorhinal cortex (pair)
stratum stellare corticis entorhinalis (par) ; lamina 2 corticis entorhinalis (par)
28463 9425
pyramidal layer of entorhinal cortex (pair) ; layer 3 of entorhinal cortex (pair)
stratum pyramidale corticis entorhinalis (par) ; lamina 3 corticis entorhinalis (par)
28466 9427
dissecting layer of entorhinal cortex (pair) ; layer 4 of entorhinal cortex (pair)
lamina dissecans corticis entorhinalis (par) ; lamina 4 corticis entorhinalis (par)
28469 9428
internal principal layer of entorhinal cortex (pair) ; layer 5 of entorhinal cortex (pair)
stratum principale internum corticis entorhinalis (par) ; lamina 5 corticis entorhinalis (par)
28472 9429
magnocellular layer of entorhinal cortex (pair) ; sublayer 5a of entorhinal cortex (pair)
stratum magnocellulare corticis entorhinalis (par) ; sublamina 5a corticis entorhinalis (par)
28475 9430
parvocellular layer of entorhinal cortex (pair) ; sublayer 5b of entorhinal cortex (pair)
stratum parvocellulare corticis entorhinalis (par) ; sublamina 5b corticis entorhinalis (par)
28478 9431
layer 5c of entorhinal cortex (pair) ; sublayer 5c of entorhinal cortex (pair)
stratum 5c corticis entorhinalis (par) ; sublamina 5c corticis entorhinalis (par)
45465 8766
layers of perirhinal cortex (pair)
strata corticis perirhinalis (par)
45468 8767
molecular layer of perirhinal cortex (pair) ; layer 1 of perirhinal cortex (pair)
stratum moleculare corticis perirhinalis (par) ; lamina 1 corticis perirhinalis (par)
45471 8768
cell island layer of perirhinal cortex (pair); layer 2 of perirhinal cortex (pair)
stratum stellare corticis perirhinalis (par) ; lamina 2 corticis perirhinalis (par)
45474 8769
external pyramidal layer of perirhinal cortex (pair) ; layer 3 of perirhinal cortex (pair)
stratum pyramidale externum corticis perirhinalis (par) ; lamina 3 corticis perirhinalis (par)
45477 8770
dissecting layer of perirhinal cortex (pair) ; layer 4 of perirhinal cortex (pair)
lamina dissecans corticis perirhinalis (par) ; lamina 4 corticis perirhinalis (par)
25047 8771
internal pyramidal layer of perirhinal cortex (pair) ; layer 5 of perirhinal cortex (pair)
stratum pyramidale internum corticis perirhinalis (par) ; lamina 5 corticis perirhinalis (par)
25054 8772
multiform layer of perirhinal cortex (pair) ; layer 6 of perirhinal cortex (pair)
stratum multiforme corticis perirhinalis (par) ; lamina 6 corticis perirhinalis (par)
28499 9454
cingulate cortex (pair)
cortex cingularis (par)
28487 9433
retrosplenial cortex (pair)
cortex retrosplenialis (par)
28490 9434
ectosplenial cortex (pair)
cortex ectosplenialis (par)
28493 9443
granular retrosplenial cortex (pair)
cortex retrosplenialis granularis (par)
28496 9453
dysgranular retrosplenial cortex (pair)
cortex retrosplenialis dysgranularis (par)
34 lines
97.6 %
100.0 %
Scientific notes
Libelle of note
The Mesocortex (Rose M 1927 Der Allocortex bei Tier und Mensch. I. Teil. J Psychol Neurol (Lpz) 34:1-11) comprises the Proisocortex, a transition area between the Isocortex and the Allocortex, and the Periallocortex, the adjoining part of the Allocortex (Filimonoff 1947); also known as Paralimbic cortex (Mesulam 1985). The Periallocortex can further be subdivided into the Peripaleocortex (Claustrum) and the Periarchicortex (Entorhinal, Presubicular and Retrosplenial cortices and part of the Cingulate gyrus; Filimonoff 1947; Zilles and Amunts 2012).
The Cortex perirhinalis (Perirhinal cortex; BA35) and the Transentorhinal subregion of Braak and Braak (1992) are somewhat synonymous terms (Augustinack JC, Huber KE, Stevens AA, et al. 2013 Predicting the location of human perirhinal cortex, Brodmann's area 35, from MRI. Neuroimage 64:32-42). In other studies (Ding S-L, Van Hoesen GW 2010 Borders, extent, and topography of human perirhinal cortex as revealed using multiple modern neuroanatomical and pathological markers. Hum Brain Mapp 31:1359-1379), BA 36 is included within the Perirhinal cortex. This is unfortunate since BA35 is periarchicortex but BA36 (Ectorhinal cortex) is truly isocortex. The Layers of BA35 are comparable to those of the adjacent Entorhinal cortex.
In the Cortex entorhinalis (Entorhinal cortex), Insausti et al. (Insausti R, Tuñón T, Sobreviela T, et al. 1995 The human entorhinal cortex: A cytoarchitectonic analysis. J Comp Neurol 355:171-198) distinguished 8 subfields (EO, ER, ELR, EMI, EI, ELC, EC and ECL). For the layers of the Entorhinal cortex, the subdivision by Insausti and Amaral (2012) into six Laminae is advocated. To avoid confusion with isocortical layers, here, arabic numerals are used as in the literature. TH Latin and English terms (H4.8.03.104/114) are added. For Layers 2 and 3, the general term External principal layer is advocated, for Layer 5 Internal principal layer, following Braak H, Braak E (1992 The human entorhinal cortex: Normal morphology and lamina-specific pathology in various diseases. Neurosci Res 15:6-31). Layer 2 is made up of islands of relatively large and darkly stained modified pyramidal and stellate cells (Braak and Braak 1992: Pre-a). Layer 3 corresponds to layers Pre-ß and Pre-? of Braak and Braak (1992). The layers Pre-a, Pre-ß and Pre-? form their External principal layer. Layer 5 corresponds to the Internal principal layer with sublayers Pri-a, Pri-ß and Pri-? of Braak and Braak (1992).
The complex Cortex retrosplenialis (Retrosplenial cortex) consists of Periarchicortical (BA26) and Proisocortical (BA29, 30) areas (Braak H 1980 Architectonics of the Human Telencephalic Cortex. Springer, Berlin-Heidelberg-New York; Zilles and Amunts 2012). The TH nomenclature (H4.8.03.122/129) seems to combine these different structures. The Cortex ectosplenialis (Ectosplenial cortex; BA26) has a primitive three-laminar pattern with Molecular, Densocellular and Multiform layers. The Cortex retrosplenialis granularis (Granular retrosplenial cortex; BA29) shows a four-layered structure: Molecular, External and Internal pyramidal and Multiform layers. The Cortex retrosplenialis dysgranularis (Dysgranular retrosplenial cortex; BA 30) shows a further progression of laminar differentiation with an additional (Internal) granular layer.
For further subdivision of the Cortex cingularis (Cingulate cortex) with Layers, see Vogt BA, Palomero-Gallager N (2012) Cingulate cortex. In: Mai JK, Paxinos G, eds: The Human Nervous System, 3rd ed. Elsevier, Amsterdam, pp 934-987.
TH subdivision into six layers (H4.8.03.115/121) suggested a well-divided structure. Insausti and Amaral (2012) emphasized that the laminar organization of the Presubiculum is complex and only poorly understood. They described a single, superficially located cellular layer made up of External and Internal principal layers. Their subdivision is followed here.
Type of list P4
List Unit Identifier 6129
Number of children 101 (validated)
Number of units 34 (validated)
Signature 700 (validated since 12.12.2020)
Date: 31.12.2021